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VTGKIDEIYRRIEQSGRSEVFIHLRPRDQVEADHASATSGPLAGLVLAVKDNVDVAGLPTTAACPGYSFV PDTDAPVVAVLKAAGAVVIGKTNLDQFATGLVGTRSPYGPVRNAQRPEYISGGSSSGSAVAVALGFADIA IGTDTAGSGRVPAGLQGIVGIKPTLGVVPNTGVVPACASYDCVTIFAADLPTAQAAMGVMATAEPHRAWP ADLMLAAPPAPRVAVPAQVPELDVAWQAAFTEAVQRAQAAGFETVPIDMTPFFAAADLLYNGALVAERWD AVGEFIESAGPGAGLDPTVSAIISAAGRHSAADLLRDRRRVEHLRDEAFAALAGCDALLVPTAPEHPRIA DVAADPIGVNARMGRFTNFCNLFDMCAIAIPAGTTSDGAQFGITLLAPAFGDAVVADLAARFLAVPAAAS WAEAATEVTEIAVFGAHLRGQPLEHELTGRGARWGGPVVTAPHYRLFALDTEPPKPGLVRVDSDGSAIEG ERWVLSPAALGEFLAELPAPMLLGKVELSDGRWVTGFGCDHAAPAHGRDITEFGGWLRAQSALSAR
Operon Prediction Model: Genebank
Paralogs
| species | id | gene | e-value | identity (len) | annotation |
| M. smegmatis MC2 155 | MSMEG_2189 | atzF | - | 100% (556) | allophanate hydrolase |
| M. smegmatis MC2 155 | MSMEG_2986 | - | 1e-43 | 37.11% (380) | amidohydrolase, AtzE family protein |
| M. smegmatis MC2 155 | MSMEG_4062 | - | 2e-37 | 33.87% (375) | glutamyl-tRNA(Gln) amidotransferase subunit A |
Closest Orthologs (e-value cutoff: 1e-4)
| species | id | gene | e-value | identity (len) | annotation |
| M. bovis AF2122 / 97 | Mb2384 | amiA2 | 3e-35 | 31.53% (425) | amidase |
| M. gilvum PYR-GCK | Mflv_2712 | - | 1e-31 | 30.42% (424) | amidase |
| M. tuberculosis H37Rv | Rv2363 | amiA2 | 3e-35 | 31.53% (425) | amidase |
| M. leprae Br4923 | MLBr_01702 | gatA | 1e-27 | 28.64% (433) | aspartyl/glutamyl-tRNA amidotransferase subunit A |
| M. abscessus ATCC 19977 | MAB_1558c | - | 0.0 | 64.23% (534) | allophanate hydrolase |
| M. marinum M | MMAR_3673 | amiA2 | 2e-33 | 31.69% (426) | amidase AmiA2 |
| M. avium 104 | MAV_2032 | - | 5e-35 | 31.82% (418) | amidase |
| M. thermoresistible (build 8) | TH_0712 | - | 1e-135 | 60.52% (423) | CONSERVED HYPOTHETICAL PROTEIN |
| M. ulcerans Agy99 | MUL_3616 | amiA2 | 1e-33 | 31.28% (422) | amidase |
| M. vanbaalenii PYR-1 | Mvan_4052 | - | 1e-171 | 58.43% (534) | amidase |
CLUSTAL 2.0.9 multiple sequence alignment
Mb2384|M.bovis_AF2122/97 MVGASGSDAGAISGSGNQRLPTLTDLLYQLATRAVTSEELVRRSLRAIDV
Rv2363|M.tuberculosis_H37Rv MVGASGSDAGAISGSGNQRLPTLTDLLYQLATRAVTSEELVRRSLRAIDV
MMAR_3673|M.marinum_M MVGASGFASGAVSGSGGPRLPTLTDLLYQLASGSVTSVELTRRSLREIEV
MUL_3616|M.ulcerans_Agy99 MVGASGFASGAVSGSGGPRLPTLTDLLYQLASGSVTSVELTRRSLREIEV
MAV_2032|M.avium_104 ----------------------------------------MGRSLHAIHA
Mflv_2712|M.gilvum_PYR-GCK MPDSS--ARSTSRNRFPDRFPTVANQLFQLASGTATSVDLVRRSLVAIEA
MAB_1558c|M.abscessus_ATCC_199 -----------------------------------------------MGQ
Mvan_4052|M.vanbaalenii_PYR-1 --------------------------------------------------
MSMEG_2189|M.smegmatis_MC2_155 ----------------------------------------MTGKIDEIYR
TH_0712|M.thermoresistible__bu -----------------------------------MSAPSAVERVHAAYA
MLBr_01702|M.leprae_Br4923 ----------MIGSLHEIIRLDASTLAAKIVAKELSSVEITQACLDQIEA
Mb2384|M.bovis_AF2122/97 SQPTLNAFRVVLTESALADAAAADKRRAAGDT----APLLGIPIAVKDDV
Rv2363|M.tuberculosis_H37Rv SQPTLNAFRVVLTESALADAAAADKRRAAGDT----APLLGIPIAVKDDV
MMAR_3673|M.marinum_M SQSTLNAFRVVLTESALADAAAADRRRAAGDA----APLLGIPIAVKDDV
MUL_3616|M.ulcerans_Agy99 SQSTLNAFRVVLTESALADAAAADRRRAAGDA----APLLGIPIAVKDDV
MAV_2032|M.avium_104 SQPTLNAFRVVLTESALADAAEADRRRAAGDT----APLLGIPIAVKDDV
Mflv_2712|M.gilvum_PYR-GCK SQPTLNAFRVVLAERALADAAEADRLRAAGHDR-ARRPLLGIPIAVKDDV
MAB_1558c|M.abscessus_ATCC_199 GTRTLLDMTVWIMRRP-----EDDIAAELEAS---SGPLAGVRLAVKDNV
Mvan_4052|M.vanbaalenii_PYR-1 -------MAVWISRRR-----GPDVAAELSESRSAAGHLAGVRLAVNDNV
MSMEG_2189|M.smegmatis_MC2_155 RIEQSGRSEVFIHLRP-----RDQVEADHASAT--SGPLAGLVLAVKDNV
TH_0712|M.thermoresistible__bu AIERVDRPEIWITLRPLADALADAVRVDAAARTGAELPLAGLVAAVKNNI
MLBr_01702|M.leprae_Br4923 TDDTYRAFLHVAAEKALSAAAAVDKAVAAGGQLS--STLAGVPLALKDVF
* *: *::: .
Mb2384|M.bovis_AF2122/97 DVAGVPTAFGTQGYVAPAT-DDCEVVRRLKAAGAVIVGKTNTCELGQWPF
Rv2363|M.tuberculosis_H37Rv DVAGVPTAFGTQGYVAPAT-DDCEVVRRLKAAGAVIVGKTNTCELGQWPF
MMAR_3673|M.marinum_M DVAGVPTAFGTQGYVRPAA-HDSEVVRRLKAAGAVIIGKTNTCELGQWPF
MUL_3616|M.ulcerans_Agy99 DVAGVPTAFGTQGYVQPAA-HDSEVVRRLKAAGAVIIGKTNTCELGQWPF
MAV_2032|M.avium_104 DIAGVPTAFGTEGSVPPAT-HDAEVVRRLKAAGAVIVGKTNTCELGQWPF
Mflv_2712|M.gilvum_PYR-GCK DVAGVPTRFGASGEVPEAV-ADAEVVRRLRAAGAVIVGKTNTCELGQWPF
MAB_1558c|M.abscessus_ATCC_199 DVGGVPTTAACPEYAYVPD-QDAPAVAALRAAGAVVVGKTNLDQFATGLV
Mvan_4052|M.vanbaalenii_PYR-1 DVAGLPTTAGCPQFAYLPD-SDAAAVAALRGAGAVVVGKTNLDQFAAGLV
MSMEG_2189|M.smegmatis_MC2_155 DVAGLPTTAACPGYSFVPD-TDAPVVAVLKAAGAVVIGKTNLDQFATGLV
TH_0712|M.thermoresistible__bu DVAGIPTTAGCPTFATCAATADAPAVARLRTAGAVVIGATNLDQFATGLV
MLBr_01702|M.leprae_Br4923 TTVDMPTTCGSKILQGWHSPYDATVTTRLRAAGIPILGKTNMDEFAMGSS
.:** . *. .. *: ** ::* ** ::.
Mb2384|M.bovis_AF2122/97 TSGPGFGHTRNPWSRRHTPGGSSGGSAAAVAAGLVTAAIGSDGAGSIRIP
Rv2363|M.tuberculosis_H37Rv TSGPGFGHTRNPWSRRHTPGGSSGGSAAAVAAGLVTAAIGSDGAGSIRIP
MMAR_3673|M.marinum_M TSGPGFGHTRNPWARRHTPGGSSGGSAAAVAAGLVTAAIGSDGAGSIRVP
MUL_3616|M.ulcerans_Agy99 TSGPGFGHTRNPWARRHTPGGSSGGSAAAVAAGLVTAAIGSDGAGSIRVP
MAV_2032|M.avium_104 TSGPGFGHTRNPWSRRHTPGGSSGGSAAAVAAGLVTAAIGSDGAGSIRIP
Mflv_2712|M.gilvum_PYR-GCK TSGPAFGHTRNPWSREHTPGGSSGGSAAAVAAGLVAAAIGSDGAGSVRIP
MAB_1558c|M.abscessus_ATCC_199 GTRSPYGAVSDSRRPEYISGGSSSGSAVAVATGQADIAIGTDTAGSGRVP
Mvan_4052|M.vanbaalenii_PYR-1 GTRSPYGAVPDSRRPEFISGGPSSGSAVAVATGEADIAIAPDATG--CIP
MSMEG_2189|M.smegmatis_MC2_155 GTRSPYGPVRNAQRPEYISGGSSSGSAVAVALGFADIAIGTDTAGSGRVP
TH_0712|M.thermoresistible__bu GTRSPYGAVRDARRPDRISGGSSSGSAVAVALGLVDVALGTDTAGSGRVP
MLBr_01702|M.leprae_Br4923 TENSAYGPTRNPWNVDRVPGGSGGGSAAALAAFQAPLAIGSDTGGSIRQP
. :* . :. .**...***.*:* . *:..* * *
Mb2384|M.bovis_AF2122/97 AAWTHLVGIKPQRGRISTWPLPEAFNGVTVNGVLARTVEDAALVLDAASG
Rv2363|M.tuberculosis_H37Rv AAWTHLVGIKPQRGRISTWPLPEAFNGVTVNGVLARTVEDAALVLDAASG
MMAR_3673|M.marinum_M AAWTHLVGIKPQRGRISTWPWPETFNGITVNGVLARTVADAALVLDAASG
MUL_3616|M.ulcerans_Agy99 AAWTHLVGIKPQRGRISTWPWPETFNGITVNGVLARTVADAALVLDAASG
MAV_2032|M.avium_104 AAWTHLVGIKPQRGRISTWPLPEAFNGITVNGVLARTVADAALVLDAASG
Mflv_2712|M.gilvum_PYR-GCK AAWTHLVGIKPQRGRISTSPLPEAFNGITVNGVLARTVSDAAMVLDAAAG
MAB_1558c|M.abscessus_ATCC_199 AGLQGIVGIKPTVGVISTEGVVPACESYDCVTIFAPSLTTANAAMAAMGA
Mvan_4052|M.vanbaalenii_PYR-1 AALQGIVAIKPTVGVISTDGAVPASESYDCITILAATLNLADTAMAVLAA
MSMEG_2189|M.smegmatis_MC2_155 AGLQGIVGIKPTLGVVPNTGVVPACASYDCVTIFAADLPTAQAAMGVMAT
TH_0712|M.thermoresistible__bu AALQGIVGIKPTIGVVPTDGVVPACRSYDCVTVFARDLDTADAAMGVLAG
MLBr_01702|M.leprae_Br4923 AALTATVGVKPTYGTVSRYGLVACASSLDQGGPCARTVLDTALLHAVIAG
*. *.:** * :. . * : : . .
Mb2384|M.bovis_AF2122/97 NVEGDRHQPPPVTVSDFVGIAPGP--------------LKIALSTHFPYT
Rv2363|M.tuberculosis_H37Rv NVEGDRHQPPPVTVSDFVGIAPGP--------------LKIALSTHFPYT
MMAR_3673|M.marinum_M NVEGDLHKPAPLTASDYVGIAPGP--------------LKIAMSTRFPYT
MUL_3616|M.ulcerans_Agy99 NAEGDLHKPAPLTASDYVGIAPGP--------------LKIAMSTRFPYT
MAV_2032|M.avium_104 NVDGDRHKPPPITASDYVGKAPGP--------------LNIALSTRFPYT
Mflv_2712|M.gilvum_PYR-GCK NAAGDLHTPAPVTVSDFVSRAPGP--------------LRVAVSTRFPFT
MAB_1558c|M.abscessus_ATCC_199 AAGPRLWPANTRLAAPPQPTIAVP--------------RELPALDEVWCN
Mvan_4052|M.vanbaalenii_PYR-1 GAPARPWPTDVKLAAPPVPVIAIP--------------DELPYLDDQWRV
MSMEG_2189|M.smegmatis_MC2_155 AEPHRAWPADLMLAAPPAPRVAVP--------------AQVPELDVAWQA
TH_0712|M.thermoresistible__bu GG--RPFPPDAPLTVPPTTRVAVP--------------GRLPGLSPAWRQ
MLBr_01702|M.leprae_Br4923 HDARDSTSVEAAVPDIVGAAKAGESGDLHGVRVGVVRQLRGEGYQSGVLA
. .
Mb2384|M.bovis_AF2122/97 GFRAKLHPEILAATQRVGDQLELLGHTVVKGNPDYGLRLSWNFLARSTAG
Rv2363|M.tuberculosis_H37Rv GFRAKLHPEILAATQRVGDQLELLGHTVVKGNPDYGLRLSWNFLARSTAG
MMAR_3673|M.marinum_M GFRAKLHPEILAATRRVGEQLQLLGHTVERGNPDYGLRLSWDFLARSTAG
MUL_3616|M.ulcerans_Agy99 GFRAKLHPEILASTRRVGEQLQLLGHTVERGNPDYGLRLSWDFLARSTAG
MAV_2032|M.avium_104 GFRPKLHPEILAATRAVGKQLELLGHTVVPGNPDYGMRLSWDFLARSTAG
Mflv_2712|M.gilvum_PYR-GCK GFRATLHPEIRSALDGVADQLSQLGHTVLNADPDYTLGMSWNFLARSTSG
MAB_1558c|M.abscessus_ATCC_199 AFRGAVDDLRAAGATIIEIDLTPFLAAAK-------LLYEGALVSERYAA
Mvan_4052|M.vanbaalenii_PYR-1 AFDAAVDTLVARGARVVSVEIEPFLTAAT-------LLHDGALVSERYAA
MSMEG_2189|M.smegmatis_MC2_155 AFTEAVQRAQAAGFETVPIDMTPFFAAAD-------LLYNGALVAERWDA
TH_0712|M.thermoresistible__bu AFESACERLAGNGIELVEVDLTPFLQAAT-------LLYDGGLVAERHAA
MLBr_01702|M.leprae_Br4923 SFQAAVEQLIALGATVSEVDCPHFDYALAAYYLILPSEVSSNLARFDAMR
.* . . : : : . :
Mb2384|M.bovis_AF2122/97 LWEWAERLG-----------DEVTLDRRTVSNLRMGHVLSQAILRSARRH
Rv2363|M.tuberculosis_H37Rv LWEWAERLG-----------DEVTLDRRTVSNLRMGHVLSQAILRSARRH
MMAR_3673|M.marinum_M LWEWAECLG-----------DGVTLDPRTVTNMRAGHALSQAILRNARHH
MUL_3616|M.ulcerans_Agy99 LWEWAECLG-----------NGVTLDPRTVTNMRAGHALSQAILRNARHH
MAV_2032|M.avium_104 LRDWEERLG-----------DGVVLDPRTVANLRLGHLLGQAILRSARRH
Mflv_2712|M.gilvum_PYR-GCK IPEWMDRLG-----------HRVTWDSRTRSNARMGRLLSQHTLRKARAR
MAB_1558c|M.abscessus_ATCC_199 VGEFIDAH------------PEASLDPTVAQIISGARDIPAHRLVHDRAE
Mvan_4052|M.vanbaalenii_PYR-1 VGEFVDGQ------------PAAALDPTVRDIITAARDVPAHRLLRDRAA
MSMEG_2189|M.smegmatis_MC2_155 VGEFIESAG-----------PGAGLDPTVSAIISAAGRHSAADLLRDRRR
TH_0712|M.thermoresistible__bu VGTFVDEHLA---------AGDPALDPTVAQIISAAGAVPATTLLRDRAR
MLBr_01702|M.leprae_Br4923 YGLRVGDDGSRSAEEVIALTRAAGFGPEVKRRIMIGAYALSAGYYDSYYS
. . .
Mb2384|M.bovis_AF2122/97 EAADQRRVGSIFDIVDVVLAPTTAQPPPMARAFDRLGSFGTDRAIIAACP
Rv2363|M.tuberculosis_H37Rv EAADQRRVGSIFDIVDVVLAPTTAQPPPMARAFDRLGSFGTDRAIIAACP
MMAR_3673|M.marinum_M EAAAQRRVGSIFDIVDVVLAPTTAQPPPLARAFDDMGGLATDRAIIAACP
MUL_3616|M.ulcerans_Agy99 EAAAQRRVGSIFDIVDVVLAPTTAQPPPLARAFDDMGGLATDRAIIAACP
MAV_2032|M.avium_104 EAADQRRVGSIFDIVDVVLAPTTAQPPPLARAFDRLSGFGTDRAMIAACP
Mflv_2712|M.gilvum_PYR-GCK EAAAQQRIGWIFNLADVVVAPTTAQPPPEAHAFDGLGGLATDRMMIRACP
MAB_1558c|M.abscessus_ATCC_199 VQRLREQAMTSLAGADALLVPTAPLHPTIEQVNANPIGVNAT-----MGT
Mvan_4052|M.vanbaalenii_PYR-1 VGRLRDRAMAALAGTDALLVPTAPVHPTIADATADPVEINSR-----LGT
MSMEG_2189|M.smegmatis_MC2_155 VEHLRDEAFAALAGCDALLVPTAPEHPRIADVAADPIGVNAR-----MGR
TH_0712|M.thermoresistible__bu LAELTASALARLGDCAALLIPTTTEHPSIAEVAADPVGVNSR-----LGT
MLBr_01702|M.leprae_Br4923 QAQKVRTLIARDLDKAYRSVDVLVSPATPTTAFSLGEKADDPLAMYLFDL
. . .
Mb2384|M.bovis_AF2122/97 STWPWNLLGWPSINVPAGFTSDG-LPIGVQLMGPANSEGMLISLAAELEA
Rv2363|M.tuberculosis_H37Rv STWPWNLLGWPSINVPAGFTSDG-LPIGVQLMGPANSEGMLISLAAELEA
MMAR_3673|M.marinum_M LTWPWNLLGWPSINVPAGFTVDG-LPIGVQLMGPAESEGLLISLAAELEA
MUL_3616|M.ulcerans_Agy99 LTWPWNLLGWPSINVPAGFTVDG-LPIGVQLMGPAESEGLLISLAAELEA
MAV_2032|M.avium_104 LTFPWNVLGWPSINVPAGFTSDG-LPIGVQLMGPANSEGMLISLAAELEA
Mflv_2712|M.gilvum_PYR-GCK VTWPWNLLGWPSINVPAGFTSDG-LPIGVQLMGPADSEPLLISLAAALEA
MAB_1558c|M.abscessus_ATCC_199 YTNFCNLFDLCAVSVPAGTAGD--AQFGVTVIARAFDDAVAFDIAALATG
Mvan_4052|M.vanbaalenii_PYR-1 YTNFCNLFDMCAIAVPAGTAGG--AQFGVTVLARAFEDAVALDVAALLVD
MSMEG_2189|M.smegmatis_MC2_155 FTNFCNLFDMCAIAIPAGTTSDG-AQFGITLLAPAFGDAVVADLAARFLA
TH_0712|M.thermoresistible__bu YTNFCNLMDLCAVAVPAGTTSDG-AQFGVTVLGRAGADALALDLARLVSA
MLBr_01702|M.leprae_Br4923 CTLPLNLAGHCGMSVPSGLSSDDDLPVGLQIMAPALADDRLYRVGAAYEA
* *: . .: :*:* : . .*: ::. * : :.
Mb2384|M.bovis_AF2122/97 VSGWAT--------------------------------------------
Rv2363|M.tuberculosis_H37Rv VSGWAT--------------------------------------------
MMAR_3673|M.marinum_M VSGWAT--------------------------------------------
MUL_3616|M.ulcerans_Agy99 VSGWAT--------------------------------------------
MAV_2032|M.avium_104 VSGWAS--------------------------------------------
Mflv_2712|M.gilvum_PYR-GCK INGWAT--------------------------------------------
MAB_1558c|M.abscessus_ATCC_199 APAGQDVWPAAITLSHELAVFG---AHLK-----GGPLEFQLTDLGARWV
Mvan_4052|M.vanbaalenii_PYR-1 EAPSASPWPLAAAEVVELVVFG---AHLR-----GGPLVHQLTDRGARFA
MSMEG_2189|M.smegmatis_MC2_155 VPAAAS-WAEAATEVTEIAVFG---AHLR-----GQPLEHELTGRGARWG
TH_0712|M.thermoresistible__bu PVDGAAPPGVPAPPTPAVPWPXXXAADLTDDDHLGRPRGDTETGQRLELL
MLBr_01702|M.leprae_Br4923 ARGPLP--------------------------------------------
Mb2384|M.bovis_AF2122/97 -----KQPQVWWTS------------------------------------
Rv2363|M.tuberculosis_H37Rv -----KQPQVWWTS------------------------------------
MMAR_3673|M.marinum_M -----KQPKVWWNTGTSTPPIHSVRPPRR---------------------
MUL_3616|M.ulcerans_Agy99 -----KQPKVWWNTGTSTPPIHSVRPPRR---------------------
MAV_2032|M.avium_104 -----KQPQPWWDQSADTPPAPGQPAPTG---------------------
Mflv_2712|M.gilvum_PYR-GCK -----LQPENWWAPQPPTEPDLTAPDSVTGQVA-----------------
MAB_1558c|M.abscessus_ATCC_199 GPIRT-ATKYRMSALRTTPPKPGLTRSEE--NGVSIGGEVWRLSPAALGT
Mvan_4052|M.vanbaalenii_PYR-1 GEITT-APRYRMTALPTTPPKPGVTRVPDGVEGAALSGHRWLLSPAALGS
MSMEG_2189|M.smegmatis_MC2_155 GPVVT-APHYRLFALDTEPPKPGLVRVDS--DGSAIEGERWVLSPAALGE
TH_0712|M.thermoresistible__bu AHLVTGGTRAPATVLAAVVHGELLALAPFGTADGVVARAASRLVTMASGL
MLBr_01702|M.leprae_Br4923 ------------SAI-----------------------------------
Mb2384|M.bovis_AF2122/97 --------------------------------------------------
Rv2363|M.tuberculosis_H37Rv --------------------------------------------------
MMAR_3673|M.marinum_M --------------------------------------------------
MUL_3616|M.ulcerans_Agy99 --------------------------------------------------
MAV_2032|M.avium_104 --------------------------------------------------
Mflv_2712|M.gilvum_PYR-GCK --------------------------------------------------
MAB_1558c|M.abscessus_ATCC_199 FLAQLPEPMLLGKVECDD-GLWRTGFGCDG-AAAQAGVDISEHGSWPAAI
Mvan_4052|M.vanbaalenii_PYR-1 FLAALPAPMQLGAVEFAD-GSWRTAFGCD---ASVQGVDISGYGSWTSAL
MSMEG_2189|M.smegmatis_MC2_155 FLAELPAPMLLGKVELSD-GRWVTGFGCDH-AAPAHGRDITEFGGWLRAQ
TH_0712|M.thermoresistible__bu DPHGLGVPEVFWMTQQRDYRAAAQGFASGTRDGLTTWLMLTARALHAGAR
MLBr_01702|M.leprae_Br4923 --------------------------------------------------
Mb2384|M.bovis_AF2122/97 -------------
Rv2363|M.tuberculosis_H37Rv -------------
MMAR_3673|M.marinum_M -------------
MUL_3616|M.ulcerans_Agy99 -------------
MAV_2032|M.avium_104 -------------
Mflv_2712|M.gilvum_PYR-GCK -------------
MAB_1558c|M.abscessus_ATCC_199 AAGAV--------
Mvan_4052|M.vanbaalenii_PYR-1 ASGAVRQT-----
MSMEG_2189|M.smegmatis_MC2_155 SALSAR-------
TH_0712|M.thermoresistible__bu EAISIAEAQAEAN
MLBr_01702|M.leprae_Br4923 -------------