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M. ulcerans Agy99 MUL_1082 (ilvD)

annotation: dihydroxy-acid dehydratase
coordinates: 1145497 - 1147191
length: 564

PEADIKPRSRDVTDGLEKAAARGMLRAVGMVDEDFAKAQIGVASSWNEITPCNLSLDRLAKSVKEGVFAA
GGYPLEFGTISVSDGISMGHEGMHFSLVSREVIADSVETVMQAERLDGSVLLAGCDKSLPGMLMAAARLD
LASVFLYAGSILPGVAKLSDGSEREVTIIDAFEAVGACSRGLMSRADVDAIERAICPGEGACGGMYTANT
MASAAEALGMSLPGSAAPPATDRRRDGFARQSGQAVIELLRQGITARDIMTREAFENAIAVVMAFGGSTN
AVLHLLAIAHEANVELTLDDFSRIGSKVPHLADVKPFGRHVMSHVDHIGGVPVVMKTLLDAGLLHGDCLT
VTGRTVAENLEAIAPPDPDGKVLRALINPIHATGGITILRGSLAPEGAVVKTAGLDTDVFEGTARVFDGE
RAALDALEDGTITHGDVVVIRYEGPKGGPGMREMLAITGAIKGAGLGKDVLLLTDGRFSGGTTGFCVGHI
APEAVDAGPIAFVRDGDRIRLDVAGRTLDVLTDPPEFAARQQNFTPPAPRYKTGVLAKYVKLVGSAAIGA
VCG*

Operon Prediction Model: Genebank

Paralogs

species id gene e-value identity (len) annotation
M. ulcerans Agy99 MUL_1082 ilvD - 100% (564) dihydroxy-acid dehydratase

Closest Orthologs (e-value cutoff: 1e-4)

species id gene e-value identity (len) annotation
M. bovis AF2122 / 97 Mb0195c ilvD 0.0 91.44% (561) dihydroxy-acid dehydratase
M. gilvum PYR-GCK Mflv_0486 - 0.0 84.97% (559) dihydroxy-acid dehydratase
M. tuberculosis H37Rv Rv0189c ilvD 0.0 91.44% (561) dihydroxy-acid dehydratase
M. leprae Br4923 MLBr_02608 ilvD 0.0 89.18% (564) dihydroxy-acid dehydratase
M. abscessus ATCC 19977 MAB_4545 - 0.0 84.70% (562) dihydroxy-acid dehydratase
M. marinum M MMAR_0432 ilvD 0.0 99.65% (564) dihydroxy-acid dehydratase IlvD
M. avium 104 MAV_4989 ilvD 0.0 91.99% (562) dihydroxy-acid dehydratase
M. smegmatis MC2 155 MSMEG_0229 ilvD 0.0 85.87% (559) dihydroxy-acid dehydratase
M. thermoresistible (build 8) TH_0983 ilvD 0.0 89.48% (561) PROBABLE DIHYDROXY-ACID DEHYDRATASE ILVD (DAD)
M. vanbaalenii PYR-1 Mvan_0173 - 0.0 87.23% (564) dihydroxy-acid dehydratase

CLUSTAL 2.0.9 multiple sequence alignment


MUL_1082|M.ulcerans_Agy99           --------------------------MPEADIKPRSRDVTDGLEKAAARG
MMAR_0432|M.marinum_M               --------------------------MPEADIKPRSRDVTDGLEKAAARG
Mb0195c|M.bovis_AF2122/97           ---------------MPQTTDEAASVSTVADIKPRSRDVTDGLEKAAARG
Rv0189c|M.tuberculosis_H37Rv        ---------------MPQTTDEAASVSTVADIKPRSRDVTDGLEKAAARG
MLBr_02608|M.leprae_Br4923          --------------------------MMPPDIKPRSRDVTDGLEKAAARG
MAV_4989|M.avium_104                ---------------MPTTDSARAADIKQPDIKPRSRDVTDGLEKAAARG
TH_0983|M.thermoresistible__bu      ------------------------MAQNQPDIKPRSRDVTDGLEKAAARG
Mvan_0173|M.vanbaalenii_PYR-1       -------------------------MPSSPDIKPRSRDVTDGLEKAAARG
Mflv_0486|M.gilvum_PYR-GCK          MWPMNRRAHNHPMPSDSRSHSLRASGSS-VDIKPRSRDVTDGLERTAARG
MAB_4545|M.abscessus_ATCC_1997      ------------MSAPHKPDSLRASGSSQPDIKPRSRDVTDGLEKTAARG
MSMEG_0229|M.smegmatis_MC2_155      ------------MPSDDKS-------TETPDIKPRSRDVTDGLEKAAARG
                                                                  **************::****

MUL_1082|M.ulcerans_Agy99           MLRAVGMVDEDFAKAQIGVASSWNEITPCNLSLDRLAKSVKEGVFAAGGY
MMAR_0432|M.marinum_M               MLRAVGMVDEDFAKAQIGVASSWNEITPCNLSLDRLAKSVKEGVFAAGGY
Mb0195c|M.bovis_AF2122/97           MLRAVGMDDEDFAKPQIGVASSWNEITPCNLSLDRLANAVKEGVFSAGGY
Rv0189c|M.tuberculosis_H37Rv        MLRAVGMDDEDFAKPQIGVASSWNEITPCNLSLDRLANAVKEGVFSAGGY
MLBr_02608|M.leprae_Br4923          MLRAVGMNDDDFAKAQIGVASSWNEITPCNLSLDRLAKAVKEGVFSAGGY
MAV_4989|M.avium_104                MLRAVGMGDEDFAKPQIGVASSWNEITPCNLSLDRLAKAVKEGVFAAGGY
TH_0983|M.thermoresistible__bu      MLRAVGMGDEDFAKPQIGVASSWNEITPCNLSLDRLAKAVKDGVHAAGGY
Mvan_0173|M.vanbaalenii_PYR-1       MLRAVGMGDEDFAKPQIGVGSSWNEITPCNLSLDRLAKAVKEGVFEAGGF
Mflv_0486|M.gilvum_PYR-GCK          MLRAVGMTDDDWVKPQIGVGSSWNEITPCNMSLQRLAQAVKGGVHEAGGY
MAB_4545|M.abscessus_ATCC_1997      MLRAVGMGDDDWVKPQIGVGSSWNEITPCNMSLQRLAHSVKDGVHEAGGY
MSMEG_0229|M.smegmatis_MC2_155      MLRAVGMGDADWAKPQIGVGSSWNEITPCNMSLQRLAKSVKDGVHEAGGY
                                    ******* * *:.*.****.**********:**:***::** **. ***:

MUL_1082|M.ulcerans_Agy99           PLEFGTISVSDGISMGHEGMHFSLVSREVIADSVETVMQAERLDGSVLLA
MMAR_0432|M.marinum_M               PLEFGTISVSDGISMGHEGMHFSLVSREVIADSVETVMQAERLDGSVLLA
Mb0195c|M.bovis_AF2122/97           PLEFGTISVSDGISMGHEGMHFSLVSREVIADSVEVVMQAERLDGSVLLA
Rv0189c|M.tuberculosis_H37Rv        PLEFGTISVSDGISMGHEGMHFSLVSREVIADSVEVVMQAERLDGSVLLA
MLBr_02608|M.leprae_Br4923          PLEFGTISVSDGISMGHQGMHFSLVSREVIADSVETVMQAERLDGSVLLA
MAV_4989|M.avium_104                PLEFGTISVSDGISMGHEGMHFSLVSREVIADSVETVMQAERLDGSVLLA
TH_0983|M.thermoresistible__bu      PLEFGTISVSDGISMGHEGMHFSLVSREVIADSVETVMQAERLDGSVLLA
Mvan_0173|M.vanbaalenii_PYR-1       PMEFGTISVSDGISMGHEGMHFSLVSREIIADSVETVMQAERLDGSVLLA
Mflv_0486|M.gilvum_PYR-GCK          PLEFGTISVSDGISMGHEGMHFSLVSREVIADSVETVVQAERLDGTVLLA
MAB_4545|M.abscessus_ATCC_1997      PLEFGTISVSDGISMGHEGMHFSLVSREVIADSVETVMQAERLDGSVLLA
MSMEG_0229|M.smegmatis_MC2_155      PLEFGTISVSDGISMGHEGMHFSLVSREVIADSVETVVQAERLDGTVLLA
                                    *:***************:**********:******.*:*******:****

MUL_1082|M.ulcerans_Agy99           GCDKSLPGMLMAAARLDLASVFLYAGSILPGVAKLSDGSEREVTIIDAFE
MMAR_0432|M.marinum_M               GCDKSLPGMLMAAARLDLASVFLYAGSILPGVAKLSDGSEREVTIIDAFE
Mb0195c|M.bovis_AF2122/97           GCDKSLPGMLMAAARLDLAAVFLYAGSILPGRAKLSDGSERDVTIIDAFE
Rv0189c|M.tuberculosis_H37Rv        GCDKSLPGMLMAAARLDLAAVFLYAGSILPGRAKLSDGSERDVTIIDAFE
MLBr_02608|M.leprae_Br4923          GCDKSLPGMLMAAARLDLASVFLYAGSILPGRTKLSDGTEHEVTLIDAFE
MAV_4989|M.avium_104                GCDKSLPGMLMAAARLDLASVFLYAGSILPGVAKLSDGSEREVTIIDAFE
TH_0983|M.thermoresistible__bu      GCDKSLPGMLMAAARLDLAAVFLYAGSILPGIAKLSDGTEREVTIIDAFE
Mvan_0173|M.vanbaalenii_PYR-1       GCDKSLPGMLMAAARLDLASVFLYAGSILPGRVKLSDGSEKDVTIIDAFE
Mflv_0486|M.gilvum_PYR-GCK          GCDKSIPGMLMAAARLDLASVFFYNGSIMPGVAKLTDGSEKEVTIIDAFE
MAB_4545|M.abscessus_ATCC_1997      GCDKSIPGMLMAAARLDLASVFLYNGSIMPGKAKLTDGTEKEVTIIDAFE
MSMEG_0229|M.smegmatis_MC2_155      GCDKSIPGMLMAAARLDLASVFFYNGSIMPGVAKLTDGTEKEVTIIDAFE
                                    *****:*************:**:* ***:** .**:**:*::**:*****

MUL_1082|M.ulcerans_Agy99           AVGACSRGLMSRADVDAIERAICPGEGACGGMYTANTMASAAEALGMSLP
MMAR_0432|M.marinum_M               AVGACSRGLMSRADVDAIERAICPGEGACGGMYTANTMASAAEALGMSLP
Mb0195c|M.bovis_AF2122/97           AVGACSRGLMSRADVDAIERAICPGEGACGGMYTANTMASAAEALGMSLP
Rv0189c|M.tuberculosis_H37Rv        AVGACSRGLMSRADVDAIERAICPGEGACGGMYTANTMASAAEALGMSLP
MLBr_02608|M.leprae_Br4923          AVGACSRGLMPRADVDAIERAICPGEGACGGMYTANTMASAAEALGMSLP
MAV_4989|M.avium_104                AVGACARGLMPREDVDAIERAICPGEGACGGMYTANTMASAAEALGMSLP
TH_0983|M.thermoresistible__bu      AVGACARGLMPREDVDAIERAICPGEGACGGMYTANTMASAAEALGMSLP
Mvan_0173|M.vanbaalenii_PYR-1       AVGACARGLMPREDVDAIERAICPGEGACGGMYTANTMASAAEALGMSLP
Mflv_0486|M.gilvum_PYR-GCK          AVGACARGLMSREDVDIIERAICPGEGACGGMYTANTMASAAEALGMSLP
MAB_4545|M.abscessus_ATCC_1997      AVGACARGLMSREDVDIIERAICPGEGACGGMYTANTMASAAEALGMSLP
MSMEG_0229|M.smegmatis_MC2_155      AVGACVRGLMSREDVDIIERAICPGEGACGGMYTANTMASAAEALGLSLP
                                    ***** ****.* *** *****************************:***

MUL_1082|M.ulcerans_Agy99           GSAAPPATDRRRDGFARQSGQAVIELLRQGITARDIMTREAFENAIAVVM
MMAR_0432|M.marinum_M               GSAAPPATDRRRDGFARQSGQAVIELLRQGITARDIMTREAFENAIAVVM
Mb0195c|M.bovis_AF2122/97           GSAAPPATDRRRDGFARRSGQAVVELLRRGITARDILTKEAFENAIAVVM
Rv0189c|M.tuberculosis_H37Rv        GSAAPPATDRRRDGFARRSGQAVVELLRRGITARDILTKEAFENAIAVVM
MLBr_02608|M.leprae_Br4923          GSAAPPATDRRRDGFARRSGQAVIELLRRGITARDILTKEAFENAIAVVM
MAV_4989|M.avium_104                GSAAPPATDRRRDGFARRSGQAVVELLRRGITARDILTKEAFENAIAVVM
TH_0983|M.thermoresistible__bu      GSAAPPATDRRRDGFARKSGQAVVELLRRGITARDILTKEAFENAIAVVM
Mvan_0173|M.vanbaalenii_PYR-1       GSAAPPATDRRRDGFARKSGMAVVELLRRGITARDILTKEAFENAIAVVM
Mflv_0486|M.gilvum_PYR-GCK          GSASPVAIDKRRDDFARRSGEAVVEMLRRGITARDILTKEAFENAIAVVM
MAB_4545|M.abscessus_ATCC_1997      GSASPVAIDKRREEYARKSGEAVVEMLRRGITARDILTKEAFENAIAVVM
MSMEG_0229|M.smegmatis_MC2_155      GSASPVAIDKRRDEFARRSGEAVVDMLRRGITARDILTKEAFENAIAVVM
                                    ***:* * *:**: :**:** **:::**:*******:*:***********

MUL_1082|M.ulcerans_Agy99           AFGGSTNAVLHLLAIAHEANVELTLDDFSRIGSKVPHLADVKPFGRHVMS
MMAR_0432|M.marinum_M               AFGGSTNAVLHLLAIAHEANVELTLDDFSRIGSKVPHLADVKPFGRHVMS
Mb0195c|M.bovis_AF2122/97           AFGGSTNAVLHLLAIAHEANVALSLQDFSRIGSGVPHLADVKPFGRHVMS
Rv0189c|M.tuberculosis_H37Rv        AFGGSTNAVLHLLAIAHEANVALSLQDFSRIGSGVPHLADVKPFGRHVMS
MLBr_02608|M.leprae_Br4923          AFGGSTNAILHLLAIAHEADVTLSLEDFSRIGFKVPHIADVKPFGRHVMF
MAV_4989|M.avium_104                AFGGSTNAVLHLLAIAHEADVALSLDDFSRIGSKVPHLADVKPFGRHVMT
TH_0983|M.thermoresistible__bu      AFGGSTNAVLHLLAIAHEANVKLTLEDFTRIGNKVPHLADVKPFGAHVMF
Mvan_0173|M.vanbaalenii_PYR-1       AFGGSTNAVLHLLAIAYEANVKLTLEDFTRVGAKVPHLADVKPFGAYVMN
Mflv_0486|M.gilvum_PYR-GCK          AFGGSTNAVLHLLAIAWEAQVELTLADFTRIGQKVPHLADVKPFGAYVMK
MAB_4545|M.abscessus_ATCC_1997      AFGGSTNAVLHLLAIAKEAEVELSLADFTRIGNKVPHLADVKPFGRHVMK
MSMEG_0229|M.smegmatis_MC2_155      AFGGSTNAVLHLLAIAREAEVELTLDDFTRVGSKVPHLADVKPFGRHVMK
                                    ********:******* **:* *:* **:*:*  ***:******* :** 

MUL_1082|M.ulcerans_Agy99           HVDHIGGVPVVMKTLLDAGLLHGDCLTVTGRTVAENLEAIAPPDPDGKVL
MMAR_0432|M.marinum_M               HVDHIGGVPVVMKTLLDAGLLHGDCLTVTGRTVAENLEAIAPPDPDGKVL
Mb0195c|M.bovis_AF2122/97           DVDHIGGVPVVMKALLDAGLLHGDCLTVTGHTMAENLAAITPPDPDGKVL
Rv0189c|M.tuberculosis_H37Rv        DVDHIGGVPVVMKALLDAGLLHGDCLTVTGHTMAENLAAITPPDPDGKVL
MLBr_02608|M.leprae_Br4923          DVDHIGGVPVVMKALLDAGLLHGDCLTVTGQTMAENLASIAPPDPDGQVI
MAV_4989|M.avium_104                DVDHIGGVPVMMKALLDAGLLNGDCLTVTGATVAQNLAAIAPPDPDGKVL
TH_0983|M.thermoresistible__bu      DVDHIGGVPVIMKTLLDAGLLHGDCLTVTGKTMAENLAHIAPPDPDGKVL
Mvan_0173|M.vanbaalenii_PYR-1       DVDRIGGVPVVMKALLDAGLLHGDCLTVTGKTMAENLAHIAPPDPDGKVL
Mflv_0486|M.gilvum_PYR-GCK          HVDEIGGVPVVMRALLDAGLLHGDCLTVTGQTMAENLAHIEPPDPDGKVL
MAB_4545|M.abscessus_ATCC_1997      DVDEIGGVPVVMRALLDAGLLHGDCLTVTGKTMAENLAHIAPPDPDGKVL
MSMEG_0229|M.smegmatis_MC2_155      HVDEIGGVPVVMRALLDAGLLNGECLTVTGKTMAENLAHIAPPDPDGKVL
                                    .**.******:*::*******:*:****** *:*:**  * ******:*:

MUL_1082|M.ulcerans_Agy99           RALINPIHATGGITILRGSLAPEGAVVKTAGLDTDVFEGTARVFDGERAA
MMAR_0432|M.marinum_M               RALNNPIHATGGITILRGSLAPEGAVVKTAGLDTDVFEGTARVFDGERAA
Mb0195c|M.bovis_AF2122/97           RALANPIHPSGGITILHGSLAPEGAVVKTAGFDSDVFEGTARVFDGERAA
Rv0189c|M.tuberculosis_H37Rv        RALANPIHPSGGITILHGSLAPEGAVVKTAGFDSDVFEGTARVFDGERAA
MLBr_02608|M.leprae_Br4923          RTLHNPIHPTGGITILRGSLAPDGAVVKTAGLDSDVFEGTARVFDGERAA
MAV_4989|M.avium_104                RALSDPLHPTGGITILRGSLAPEGAVVKSAGFDSDVFEGTARVFDGERAA
TH_0983|M.thermoresistible__bu      RALSNPIHPTGGITILHGSLAPEGAVVKSAGFDTSVFEGTARVFDRERAA
Mvan_0173|M.vanbaalenii_PYR-1       RALSRPIHPTGGITILHGSLAPEGAVVKSAGFDSDVFEGTARVFERERAA
Mflv_0486|M.gilvum_PYR-GCK          RAMNNPIHPTGGITILHGSLAPEGAVVKSAGFDSDVFEGTARVFERERAA
MAB_4545|M.abscessus_ATCC_1997      RAMNDPIHPTGGITILHGSLAPEGAVVKSAGFESDVFEGTARVFERERAA
MSMEG_0229|M.smegmatis_MC2_155      RAMNNPIHPTGGITILHGSLAPEGAVVKSAGFDSDVFEGTARVFDRERAA
                                    *::  *:*.:******:*****:*****:**:::.*********: ****

MUL_1082|M.ulcerans_Agy99           LDALEDGTITHGDVVVIRYEGPKGGPGMREMLAITGAIKGAGLGKDVLLL
MMAR_0432|M.marinum_M               LDALEDGTITHGDVVVIRYEGPKGGPGMREMLAITGAIKGAGLGKDVLLL
Mb0195c|M.bovis_AF2122/97           LDALEDGTITVGDAVVIRYEGPKGGPGMREMLAITGAIKGAGLGKDVLLL
Rv0189c|M.tuberculosis_H37Rv        LDALEDGTITVGDAVVIRYEGPKGGPGMREMLAITGAIKGAGLGKDVLLL
MLBr_02608|M.leprae_Br4923          LDALKDGTIAKGDVVVIRYEGPKGGPGMREMLAITGAIKGAGLGKDVLLL
MAV_4989|M.avium_104                LDALEDGTITKGDAVVIRYEGPKGGPGMREMLAITGAIKGAGLGKDVLLL
TH_0983|M.thermoresistible__bu      MDALEDGTIQAGDVVVIRYEGPKGGPGMREMLAITGAIKGAGLGKDVLLL
Mvan_0173|M.vanbaalenii_PYR-1       LDALEDGTITHGDVVVIRYEGPKGGPGMREMLAITGAIKGAGLGKDVLLM
Mflv_0486|M.gilvum_PYR-GCK          LDALEDGTITHGDVVVIRYEGPKGGPGMREMLAITGAIKGAGLGKDVLLM
MAB_4545|M.abscessus_ATCC_1997      LDALEDGTITHGDVVVIRYEGPKGGPGMREMLAITGAIKGAGLGKDVLLM
MSMEG_0229|M.smegmatis_MC2_155      MDALEDGTIQAGDVVVIRYEGPKGGPGMREMLAITGAIKGAGLGKDVLLM
                                    :***:****  **.***********************************:

MUL_1082|M.ulcerans_Agy99           TDGRFSGGTTGFCVGHIAPEAVDAGPIAFVRDGDRIRLDVAGRTLDVLTD
MMAR_0432|M.marinum_M               TDGRFSGGTTGFCVGHIAPEAVDAGPIAFVRDGDRIRLDVAGRTLDVLTD
Mb0195c|M.bovis_AF2122/97           TDGRFSGGTTGLCVGHIAPEAVDGGPIALLRNGDRIRLDVAGRVLDVLAD
Rv0189c|M.tuberculosis_H37Rv        TDGRFSGGTTGLCVGHIAPEAVDGGPIALLRNGDRIRLDVAGRVLDVLAD
MLBr_02608|M.leprae_Br4923          TDGRFSGGTTGFCVGHIAPEAVEAGPIAFLRDGDRVLLDVVGCSLDVLVD
MAV_4989|M.avium_104                TDGRFSGGTTGLCVGHIAPEAVDAGPIAFLRDGDRIRLDVANRVLDVLVD
TH_0983|M.thermoresistible__bu      TDGRFSGGTTGLCVGHVAPEAVDGGPIAFLRDGDRIRLDVAKGTLDVLVD
Mvan_0173|M.vanbaalenii_PYR-1       TDGRFSGGTTGLCVGHIAPEAVDGGPIAFIKDGDKIRLDVAKGTLDVVVD
Mflv_0486|M.gilvum_PYR-GCK          TDGRFSGGTTGLCVGHIAPEAVDGGPIAFVRDGDRIRLDVANGTLDILVD
MAB_4545|M.abscessus_ATCC_1997      TDGRFSGGTTGLCVGHIAPEAVDGGPIAFVRDGDRIRLDVANGKLDLLVD
MSMEG_0229|M.smegmatis_MC2_155      TDGRFSGGTTGLCVGHIAPEAVDGGPIAFVRDGDRIRLDVAKGTLDVLVD
                                    ***********:****:*****:.****::::**:: ***.   **::.*

MUL_1082|M.ulcerans_Agy99           PPEFAARQQNFTPPAPRYKTGVLAKYVKLVGSAAIGAVCG
MMAR_0432|M.marinum_M               PAEFAARQQNFTPPAPRYKTGVLAKYVKLVGSAAIGAVCG
Mb0195c|M.bovis_AF2122/97           PAEFASRQQDFSPPPPRYTTGVLSKYVKLVSSAAVGAVCG
Rv0189c|M.tuberculosis_H37Rv        PAEFASRQQDFSPPPPRYTTGVLSKYVKLVSSAAVGAVCG
MLBr_02608|M.leprae_Br4923          PVEFSSRKKDFIPPAPRYTTGVLAKYVKLVSSATVGAVCG
MAV_4989|M.avium_104                PAEFDSRRTAFTPPPPRYKTGVLAKYVKLVGSAAIGAVCG
TH_0983|M.thermoresistible__bu      PAEFEARKKDWEPLPPRYTTGVLAKYTKLVGSAATGAVCT
Mvan_0173|M.vanbaalenii_PYR-1       KDEFEARKVGFEPLPPVYTTGVLAKYTKLVGSAATGAVCT
Mflv_0486|M.gilvum_PYR-GCK          EAEFESRKAGFEPLPPVYTTGVLAKYTKLVGSAAVGAVCS
MAB_4545|M.abscessus_ATCC_1997      EAEIAERRKGFEPLPPRYKTGVLAKYTKLVQSAAVGAVCG
MSMEG_0229|M.smegmatis_MC2_155      EEEFEARKAGFEPLPPRYKTGVLAKYTKLVQSAAVGAVCT
                                      *:  *:  : * .* *.****:**.*** **: ****

species	id	gene	e-value	identity (len)	annotation
M. ulcerans Agy99	MUL_1082	ilvD	-	100% (564)	dihydroxy-acid dehydratase