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MTFAPSTAAASRPAGGDPAAELRIAELTDSAAVAHYIARGCLADAPLGRVGLEVEAHCHDPQDPCRRPGW DEIAAVLDALPALPGGSRVTVEPGGAVELSGPPADGVLAAIDAMRRDQAVLRPAFAEAGLGLVFLGADPL RPPQRINPGARYRAMERFFATSRSGEAGAAMMTSTASIQVNVDAGPREGWAARVRLAHALGPTMIAMTAN SPMLDGDFTGWVSSRQRVWGRMDSARCGPILGASGDDPGTDWARYALKAPVMLVHTPDAEAVTRWVPFAD WVDGRVLLGGRRPTVADLEYHLTTLFPPVRPRQWLEIRYLDSVPDTLWPAVVFTLVALLDDPVAAGLAAR AVEPVATAWDVAARVGLRDRRLYAAANACLAIAAERAPAELAEPMRRLVRLVEQGRCPGDDFADQVIEHG IAASVSRAARGPQGGP
Operon Prediction Model: Genebank
Paralogs
| species | id | gene | e-value | identity (len) | annotation |
| M. avium 104 | MAV_0399 | - | - | 100% (436) | putative glutamate--cysteine ligase |
Closest Orthologs (e-value cutoff: 1e-4)
| species | id | gene | e-value | identity (len) | annotation |
| M. bovis AF2122 / 97 | Mb3730c | gshA | 0.0 | 73.73% (434) | glutamate--cysteine ligase gshA (gamma-glutamylcysteine |
| M. gilvum PYR-GCK | Mflv_1315 | - | 1e-167 | 69.40% (415) | glutamate--cysteine ligase, GCS2 |
| M. tuberculosis H37Rv | Rv3704c | gshA | 0.0 | 73.73% (434) | glutamate--cysteine ligase gshA (gamma-glutamylcysteine |
| M. leprae Br4923 | - | - | - | - | - |
| M. abscessus ATCC 19977 | MAB_0369 | - | 1e-115 | 53.28% (396) | glutamate--cysteine ligase GshA |
| M. marinum M | MMAR_5215 | gshA | 0.0 | 73.44% (433) | glutamate--cysteine ligase GshA |
| M. smegmatis MC2 155 | MSMEG_6250 | - | 1e-169 | 67.78% (419) | glutamate--cysteine ligase |
| M. thermoresistible (build 8) | TH_0772 | gshA | 1e-160 | 67.72% (412) | GLUTAMATE--CYSTEINE LIGASE GSHA (GAMMA-GLUTAMYLCYSTEINE |
| M. ulcerans Agy99 | MUL_4094 | gcs2 | 2e-05 | 31.58% (133) | glutamate--cysteine ligase, Gcs2 |
| M. vanbaalenii PYR-1 | Mvan_5486 | - | 1e-169 | 69.64% (415) | glutamate--cysteine ligase, GCS2 |
CLUSTAL 2.0.9 multiple sequence alignment
Mflv_1315|M.gilvum_PYR-GCK --------------------------------------------------
Mvan_5486|M.vanbaalenii_PYR-1 --------------------------------------------------
MSMEG_6250|M.smegmatis_MC2_155 --------------------------------------------------
TH_0772|M.thermoresistible__bu --------------------------------------------------
Mb3730c|M.bovis_AF2122/97 --------------------------------------------------
Rv3704c|M.tuberculosis_H37Rv --------------------------------------------------
MAV_0399|M.avium_104 --------------------------------------------------
MMAR_5215|M.marinum_M --------------------------------------------------
MAB_0369|M.abscessus_ATCC_1997 --------------------------------------------------
MUL_4094|M.ulcerans_Agy99 MGVEEEFHLVDLRSRRLTTRAPDLLAKLPEDYVAELQSCVVETNSGIVDS
Mflv_1315|M.gilvum_PYR-GCK --------------------------------------------------
Mvan_5486|M.vanbaalenii_PYR-1 --------------------------------------------------
MSMEG_6250|M.smegmatis_MC2_155 --------------------------------------------------
TH_0772|M.thermoresistible__bu --------------------------------------------------
Mb3730c|M.bovis_AF2122/97 --------------------------------------------------
Rv3704c|M.tuberculosis_H37Rv --------------------------------------------------
MAV_0399|M.avium_104 --------------------------------------------------
MMAR_5215|M.marinum_M --------------------------------------------------
MAB_0369|M.abscessus_ATCC_1997 --------------------------------------------------
MUL_4094|M.ulcerans_Agy99 LDALRADLLRHRRLLVDAADEIGVGVVAAGAVPLSVPAELQITQTPRYLH
Mflv_1315|M.gilvum_PYR-GCK --------------------------------------------------
Mvan_5486|M.vanbaalenii_PYR-1 --------------------------------------------------
MSMEG_6250|M.smegmatis_MC2_155 --------------------------------------------------
TH_0772|M.thermoresistible__bu --------------------------------------------------
Mb3730c|M.bovis_AF2122/97 --------------------------------------------------
Rv3704c|M.tuberculosis_H37Rv --------------------------------------------------
MAV_0399|M.avium_104 --------------------------------------------------
MMAR_5215|M.marinum_M --------------------------------------------------
MAB_0369|M.abscessus_ATCC_1997 --------------------------------------------------
MUL_4094|M.ulcerans_Agy99 MLADYQLLAREQLICATHVHVGIDDRDEAIAIANRLLPCLPTLLALSASS
Mflv_1315|M.gilvum_PYR-GCK -----------------------------------MNLSVLSGRWVKTQR
Mvan_5486|M.vanbaalenii_PYR-1 --------------------------------------------------
MSMEG_6250|M.smegmatis_MC2_155 --------------------------------------------------
TH_0772|M.thermoresistible__bu --------------------------------------------------
Mb3730c|M.bovis_AF2122/97 --------------------------------------------------
Rv3704c|M.tuberculosis_H37Rv --------------------------------------------------
MAV_0399|M.avium_104 --------------------------------------------------
MMAR_5215|M.marinum_M --------------------------------------------------
MAB_0369|M.abscessus_ATCC_1997 --------------------------------------------------
MUL_4094|M.ulcerans_Agy99 PFWADGSDTGYASMRTLVWRRWPTTGLAAPVQSATEYDALVKALIASQVI
Mflv_1315|M.gilvum_PYR-GCK PFHTRRGWLMAHTMASDGVVEHG---------VVPLTSAEDAAQHIVDTC
Mvan_5486|M.vanbaalenii_PYR-1 ---------MACTMTSDGVVEQR---------FDPLSSAEAAAQHIVDGC
MSMEG_6250|M.smegmatis_MC2_155 ---------MALPARSDSGCAVP---------VE-FTSAEQAAAHIGANS
TH_0772|M.thermoresistible__bu ---------MTLPVTAEAPAT--------------VDSARRAAAHIEASC
Mb3730c|M.bovis_AF2122/97 ---------MTLAAMTAAASQLD----NAAPDDVEITDSSAAAEYIADGC
Rv3704c|M.tuberculosis_H37Rv ---------MTLAAMTAAASQLD----NAAPDDVEITDSSAAAEYIADGC
MAV_0399|M.avium_104 ---------MTFAPSTAAASRPAGGDPAAELRIAELTDSAAVAHYIARGC
MMAR_5215|M.marinum_M ---------MTFAPSPAATSQLD----SSRAADADIAGSAVAAQYIANGC
MAB_0369|M.abscessus_ATCC_1997 ---------MATTATEADTGR-------------TLSGHDEAAEHIARQA
MUL_4094|M.ulcerans_Agy99 TDYGMIYFDLRPSSHAPTLELRVCDSCPSVDTIVLVAGLYRAAVAREAEA
: . . . .* .
Mflv_1315|M.gilvum_PYR-GCK LKDVP----------------PGPVGLEIEAHCYDLSDPFRRPGWDELTA
Mvan_5486|M.vanbaalenii_PYR-1 LIDGP----------------VGQVGLEVEAHCFDLTDPSRRPGWDELTA
MSMEG_6250|M.smegmatis_MC2_155 LQDGP----------------IGRVGLEIEAHCFDLSNPTRRPSWDELSA
TH_0772|M.thermoresistible__bu LRDGP----------------VGRVGLEIEAHCFDVADPGRRPGWAELTD
Mb3730c|M.bovis_AF2122/97 LVDGP----------------LGRVGLEMEAHCFDPADPFRRPSWEEITE
Rv3704c|M.tuberculosis_H37Rv LVDGP----------------LGRVGLEMEAHCFDPADPFRRPSWEEITE
MAV_0399|M.avium_104 LADAP----------------LGRVGLEVEAHCHDPQDPCRRPGWDEIAA
MMAR_5215|M.marinum_M LVDGP----------------IGRVGLEIEAHCYDPADPHRRPSWEEIGD
MAB_0369|M.abscessus_ATCC_1997 FADAQ----------------VGAVGLELESHTVELTAPHRRITWNRLRE
MUL_4094|M.ulcerans_Agy99 FRAGTPAPAFPSTVGSAALWRAARSGLEGDLVDLTGPVPASRPAGDVVND
: . *** : * * :
Mflv_1315|M.gilvum_PYR-GCK VIADLPRLPGGS--------------------------------------
Mvan_5486|M.vanbaalenii_PYR-1 TITALPPLPGGS--------------------------------------
MSMEG_6250|M.smegmatis_MC2_155 VIADVPPLPGGS--------------------------------------
TH_0772|M.thermoresistible__bu VIATVGELPGGS--------------------------------------
Mb3730c|M.bovis_AF2122/97 VLEWLSPLPGGS--------------------------------------
Rv3704c|M.tuberculosis_H37Rv VLEWLSPLPGGS--------------------------------------
MAV_0399|M.avium_104 VLDALPALPGGS--------------------------------------
MMAR_5215|M.marinum_M VLEGLGKPPGGS--------------------------------------
MAB_0369|M.abscessus_ATCC_1997 VGDSVPDLPGSS--------------------------------------
MUL_4094|M.ulcerans_Agy99 LVSSLRPQLEASGDWEMIGELTRQTLLSGTSSARQRRALRRRGRLTDVVD
: .*
Mflv_1315|M.gilvum_PYR-GCK LVTVEPGGAVELS--------------------GPPATGTADAVSAMVTD
Mvan_5486|M.vanbaalenii_PYR-1 VVTVEPGGAVELS--------------------GPPLAEPASAISAMAAD
MSMEG_6250|M.smegmatis_MC2_155 RITVEPGGAVELS--------------------GPPYDGPLAAVAALQAD
TH_0772|M.thermoresistible__bu AISVEPGGAVELS--------------------GPPLDGPVAAATAMLTD
Mb3730c|M.bovis_AF2122/97 VVSVEPGGAVELS--------------------GPPADGVLAAIGAMTRD
Rv3704c|M.tuberculosis_H37Rv VVSVEPGGAVELS--------------------GPPADGVLAAIGAMTRD
MAV_0399|M.avium_104 RVTVEPGGAVELS--------------------GPPADGVLAAIDAMRRD
MMAR_5215|M.marinum_M AVTVEPGGAVELS--------------------GPPCDGILAAIDAMHRD
MAB_0369|M.abscessus_ATCC_1997 AITFEPGGAVELS--------------------GPPCSDVWSAISSMRAD
MUL_4094|M.ulcerans_Agy99 QLIAETAGRVKPTTAAVSHDGGLLAPYQLTGEAGKPADGMFASYDEAVDA
: *..* *: : * * :
Mflv_1315|M.gilvum_PYR-GCK RAVLRDVFAGAGLGLVLLGADPLRPAGRVNPGSRYAAMERFFAESGTPEA
Mvan_5486|M.vanbaalenii_PYR-1 HTVLRAAFARAGLGLVLVGADPLRPAARVNPGPRYQAMEAFFRASDSGDA
MSMEG_6250|M.smegmatis_MC2_155 RAVLRAEFARRNLGLVLLGTDPLRPTRRVNPGARYSAMEQFFTASGTAEA
TH_0772|M.thermoresistible__bu QAVLRAAFARAGYALVMLGADPVRQPQRVNPGSRYRAMETFFAASGTAAA
Mb3730c|M.bovis_AF2122/97 QAVLRSALANAGLGLVFLGADPLRSPVRVNPGARYRAMEQFFAASHSGVP
Rv3704c|M.tuberculosis_H37Rv QAVLRSALANAGLGLVFLGADPLRSPVRVNPGARYRAMEQFFAASHSGVP
MAV_0399|M.avium_104 QAVLRPAFAEAGLGLVFLGADPLRPPQRINPGARYRAMERFFATSRSGEA
MMAR_5215|M.marinum_M QAVLRSAFADAGLGLVCLGADPIRPTMRINPGARYQAMEQFFTASQSGMA
MAB_0369|M.abscessus_ATCC_1997 HEILTAVYRGAGIALASLGTDPLRRPERVNPGARYVAMERHFGAAGYGET
MUL_4094|M.ulcerans_Agy99 DGRARPNYKTALKTIEGLGVDVLRARDRDIEQERSAGNVTFRAAGHSRVQ
: :*.* :* * * . . .
Mflv_1315|M.gilvum_PYR-GCK GAAMMTSTASVQVNLEAGHRAGWADRVRLAHALG------PTMIAIAANS
Mvan_5486|M.vanbaalenii_PYR-1 GAAMMTSTASVQVNLEAGPRAGWADRVRLAHALG------PTMIAIAANS
MSMEG_6250|M.smegmatis_MC2_155 GAAMMTATASVQVNLDAGPRDGWAERVRLAHALG------PTMIAITANS
TH_0772|M.thermoresistible__bu GSAMMTSTASIQVNVEAGPRTGWAERIRLAHALG------PVMVAIAANS
Mb3730c|M.bovis_AF2122/97 GAAMMTSTAAIQVNLDAGPQEGWAERVRLAHALG------PTMIAIAANS
Rv3704c|M.tuberculosis_H37Rv GAAMMTSTAAIQVNLDAGPQEGWAERVRLAHALG------PTMIAIAANS
MAV_0399|M.avium_104 GAAMMTSTASIQVNVDAGPREGWAARVRLAHALG------PTMIAMTANS
MMAR_5215|M.marinum_M GAAMMTSTASIQVNLDAGPQSGWASRVRLAHALG------PTMIAIAANS
MAB_0369|M.abscessus_ATCC_1997 ALQMMTCTASLQVNVQSGTPRQWRDRFVLAQRIG------PTMAALSASS
MUL_4094|M.ulcerans_Agy99 VNPLDLVPRIVTADEWAQLSQGLAQRARALDAFLRDVYSEQAILADGVLS
: . : .: : * . : .: * . *
Mflv_1315|M.gilvum_PYR-GCK PVLAGKFTGWHST--------------------RQLVWSQLDSARCGP--
Mvan_5486|M.vanbaalenii_PYR-1 PVLGGDFTGWQST--------------------RQLVWSRLDSARCGP--
MSMEG_6250|M.smegmatis_MC2_155 PMLGGQFTGWCST--------------------RQRVWGQLDSARCGP--
TH_0772|M.thermoresistible__bu PLLGGRFSGWRSS--------------------RQRIWGQLDSARCGP--
Mb3730c|M.bovis_AF2122/97 PMLGGRFSGWQST--------------------RQRVWGQMDSARCGP--
Rv3704c|M.tuberculosis_H37Rv PMLGGRFSGWQST--------------------RQRVWGQMDSARCGP--
MAV_0399|M.avium_104 PMLDGDFTGWVSS--------------------RQRVWGRMDSARCGP--
MMAR_5215|M.marinum_M PMLGGEFSGWRST--------------------RQRVWGQMDDGRCGP--
MAB_0369|M.abscessus_ATCC_1997 PMLTGRPTGRRNT--------------------RQWIWDNLDPRRCAP--
MUL_4094|M.ulcerans_Agy99 AQVLDRSPGFRSTGRLAGDGVRAHISGIDLVCDRAGNWMVLEDNLRIPSG
. : . .* .: * * :: *
Mflv_1315|M.gilvum_PYR-GCK ---------ILG-VDGD-DPASDWARYALRAPVMLV--------HTP---
Mvan_5486|M.vanbaalenii_PYR-1 ---------ILG-ASGD-DPASDWARYALRAPVMLV--------HSP---
MSMEG_6250|M.smegmatis_MC2_155 ---------VLG-VDGD-DPASEWARYALRAPVMLV--------NSP---
TH_0772|M.thermoresistible__bu ---------ILGGRDGVRTPGGDWARYALRAPVMLV--------NGAGDR
Mb3730c|M.bovis_AF2122/97 ---------ILG-ASGD-HPGIDWAKYALKAPVMMV--------RSPDTQ
Rv3704c|M.tuberculosis_H37Rv ---------ILG-ASGD-HPGIDWAKYALKAPVMMV--------RSPDTQ
MAV_0399|M.avium_104 ---------ILG-ASGD-DPGTDWARYALKAPVMLV--------HTP---
MMAR_5215|M.marinum_M ---------ILG-ASAD-DPATDWAHYALKAPVMLV--------HNP---
MAB_0369|M.abscessus_ATCC_1997 ---------VEI----GVDPTESWVEYALRAPVMLV--------RNKD--
MUL_4094|M.ulcerans_Agy99 VAYAIVNRRLLGKYLPELPPPGGVADLDRVAHLLLETLRAASPPHTPDQP
: * . * ::: .
Mflv_1315|M.gilvum_PYR-GCK EAVPVTQWVPFADWADG---RVLLDDRRPTLADLDYHLTTLFPP------
Mvan_5486|M.vanbaalenii_PYR-1 DPTPVTQWVPFADWADG---RVLLDDRPPTLADLDYHLTTLFPP------
MSMEG_6250|M.smegmatis_MC2_155 DAVPVTNWVPFADWADG---RAVLGGRRPTEADLDYHLTTLFPP------
TH_0772|M.thermoresistible__bu TAEPVTDWVPFADWADG---RTALGGRRPTVADLEYHLTTLFPP------
Mb3730c|M.bovis_AF2122/97 DTRAVTDYVPFTDWVDG---RVLLDGRRATVADLVYHLTTLFPP------
Rv3704c|M.tuberculosis_H37Rv DTRAVTDYVPFTDWVDG---RVLLDGRRATVADLVYHLTTLFPP------
MAV_0399|M.avium_104 DAEAVTRWVPFADWVDG---RVLLGGRRPTVADLEYHLTTLFPP------
MMAR_5215|M.marinum_M DAVAVTHHVPFADWVDG---RVLLGDRRPTYADLDYHLTTLFPP------
MAB_0369|M.abscessus_ATCC_1997 GADAVVTRVPFQSWVDG---TMPLAGRAPTTEDLDYHLTTLFPP------
MUL_4094|M.ulcerans_Agy99 TVTLLSAGRDDPGWFEHGFLAEEMGVALVAPSDLSVRDRRVFRHGGSGGS
: .* : : : ** : :*
Mflv_1315|M.gilvum_PYR-GCK ----VRPRGFLEVRYLDSVPD---------AVWPAVVFTLSTLLDDPVAA
Mvan_5486|M.vanbaalenii_PYR-1 ----VRPRGFLEVRYLDSVPD---------ATWPAVVYTLATLLDDPAAA
MSMEG_6250|M.smegmatis_MC2_155 ----VRPRRWLEIRYLDSVPD---------ALWPAAVFTLTTLLDDPVAA
TH_0772|M.thermoresistible__bu ----VRPRGWLEIRYLDCVPD---------ALWPAVVFTLVTLLDDPSAA
Mb3730c|M.bovis_AF2122/97 ----VRPRQWLEIRYLDSVPD---------EVWPAVVFTLVTLLDDPVAA
Rv3704c|M.tuberculosis_H37Rv ----VRPRQWLEIRYLDSVPD---------EVWPAVVFTLVTLLDDPVAA
MAV_0399|M.avium_104 ----VRPRQWLEIRYLDSVPD---------TLWPAVVFTLVALLDDPVAA
MMAR_5215|M.marinum_M ----VRPRQWLEIRYLDSLSD---------VFWPAVVFTLVTLLDDPGAA
MAB_0369|M.abscessus_ATCC_1997 ----VRPRRWLELRYLDAAPD---------WWWPALAFTAVAALDDPQVA
MUL_4094|M.ulcerans_Agy99 PVDVIYSRMYEDMLLSSTGHDGAPLRSGLLEALDAGNLTIVNALGNGVAE
: .* : :: . * * * *.: .
Mflv_1315|M.gilvum_PYR-GCK DVAAEATEPVATSWDRAAQLGLDDRRLHTAAVR-----------------
Mvan_5486|M.vanbaalenii_PYR-1 DVAAEATEEVATAWDRAAQLGLRDRRLHAAATR-----------------
MSMEG_6250|M.smegmatis_MC2_155 ESAAEATRPVATAWDRAARMGLTDRHLHTAALT-----------------
TH_0772|M.thermoresistible__bu DIAAEATEPVSSAWSRAARDGLADRRLFTAANR-----------------
Mb3730c|M.bovis_AF2122/97 DLAVDAVEPVATAWDTAARIGLADRRLYLAANR-----------------
Rv3704c|M.tuberculosis_H37Rv DLAVDAVEPVATAWDTAARIGLADRRLYLAANR-----------------
MAV_0399|M.avium_104 GLAARAVEPVATAWDVAARVGLRDRRLYAAANA-----------------
MMAR_5215|M.marinum_M ELAFEAVQPVATEWDTAARAGLGDPRLHAAASR-----------------
MAB_0369|M.abscessus_ATCC_1997 DIAAEIAEPVGNAWGVAARIGLEDPALHAAAHR-----------------
MUL_4094|M.ulcerans_Agy99 DKAVYPFVPAMIEYYLGEKPALAQVPTCVCAEREQRDYVLDHLGELVVKP
* . : . : .* : .*
Mflv_1315|M.gilvum_PYR-GCK --------------CVYAAADRAPAALVESMELLTRSVEQGRSPGDDFAD
Mvan_5486|M.vanbaalenii_PYR-1 --------------CVQAAADRAPRALEESMGHLVRSVEQGRSPADDFAD
MSMEG_6250|M.smegmatis_MC2_155 --------------CVRLAAERAPAELEESMTLLMRSVQQRRSPADDFSD
TH_0772|M.thermoresistible__bu --------------CVRTAAERTPAPLLESMERLVRSVERGRCPADDFAD
Mb3730c|M.bovis_AF2122/97 --------------CLAIAARRVPTELIGAMQRLVDHVDRGVCPADDFSD
Rv3704c|M.tuberculosis_H37Rv --------------CLAIAARRVPTELIGAMQRLVDHVDRGVCPADDFSD
MAV_0399|M.avium_104 --------------CLAIAAERAPAELAEPMRRLVRLVEQGRCPGDDFAD
MMAR_5215|M.marinum_M --------------CVAIAAERAPAELADALQSLLAYVQRGRCPGDDFGD
MAB_0369|M.abscessus_ATCC_1997 --------------LVSAACAVAPPELATDMEFLLERVEQGRCPADDFID
MUL_4094|M.ulcerans_Agy99 TDGLSGSGVLIGPEATDAAIEARRRELLIQPERFVAQQLVALSTHPTFAD
* * : .. * *
Mflv_1315|M.gilvum_PYR-GCK R----------------------VVNDGIAAAVTRLAQGDM---------
Mvan_5486|M.vanbaalenii_PYR-1 R----------------------VVAHGIADAVTELAQGEL---------
MSMEG_6250|M.smegmatis_MC2_155 R----------------------VVARGIAAAVRELAKGEL---------
TH_0772|M.thermoresistible__bu L----------------------VVRRGIGDAVDRHGRGDW---------
Mb3730c|M.bovis_AF2122/97 R----------------------VIAGGIASAVTGMMHGAS---------
Rv3704c|M.tuberculosis_H37Rv R----------------------VIAGGIASAVTGMMHGAS---------
MAV_0399|M.avium_104 Q----------------------VIEHGIAASVSRAARGPQGGP------
MMAR_5215|M.marinum_M Q----------------------VTEHGVAPAIARLARGEQ---------
MAB_0369|M.abscessus_ATCC_1997 N----------------------VTEYGVEKAFSGAIG------------
MUL_4094|M.ulcerans_Agy99 DGLYPHHVDLRAFVHLRQAPRGPVTAKVLPAALTRVASRGSRIANSSSGA
* : :.
Mflv_1315|M.gilvum_PYR-GCK -----------
Mvan_5486|M.vanbaalenii_PYR-1 -----------
MSMEG_6250|M.smegmatis_MC2_155 -----------
TH_0772|M.thermoresistible__bu -----------
Mb3730c|M.bovis_AF2122/97 -----------
Rv3704c|M.tuberculosis_H37Rv -----------
MAV_0399|M.avium_104 -----------
MMAR_5215|M.marinum_M -----------
MAB_0369|M.abscessus_ATCC_1997 -----------
MUL_4094|M.ulcerans_Agy99 GSKDTWIFTDG